\[ **BT: [[eukaryotes]]** ]
---
# fungi
![[Sheldrake, Entangled Life, 2021#^fd10db]]
[^1]
[[There are few places fungi haven't colonised. Their habitats range from the sea floor to deserts, from Antarctica to our own human guts and orifices.]][^2]
[[Without fungi, no life on land would exist.]][^3]
## fungi are old as balls
#### prototaxites
[[Prototaxites were ‘living spires … scattered across the landscape’ 400 million years ago. Some were taller than a two-storey building, and were around for at least 40 million years. They are thought to have been enormous fungi.]][^4]
- see also: [[Earth#the Devonian period]]
## characteristics of fungi
### mushrooms
[[Mushrooms are the fruiting bodies of fungi (the place where spores are produced). Like the fruits of plants, they are merely one part of a much larger structure.]][^5]
[[The same way some plants use fruit, mushrooms are a fungus's way to entreat the rest of the world to help disperse their spores.]][^6]
![[Sheldrake, Entangled Life, 2021#^f202ea]]
[^7]
[[Not all fungi use the mushroom method to disperse their spores. In fact, mushroom-producing fungi are in the minority.]][^8]
#### hyphae
[[Mushrooms are made of (and by) hyphae.]][^9]
## classification of fungi
| FUNGI | Class | Order | Family | Genus | Species |
| ----------------------------- | ---------------------------- | -------------------------------------- | ------------------------------------------------------------------------------ | ---------------------------------------------------------- | ------------------------------------------------------------------------------------ |
| [[#Ascomyota (Division)]] | Pezizomycetes | Pezizales | Tuberaceae | *[[#Tuber (Genus)\|Tuber]]* | |
| | [[#Sordariomycetes (Class)]] | [[hypocrealeans\|Hypocreales (Order)]] | [[hypocrealeans#Ophiocordycipitaceae (Family)\|Ophiocordycipitaceae (Family)]] | *[[hypocrealeans#Ophiocordyceps (Genus)\|Ophiocordyceps]]* | *[[hypocrealeans#Ophiocordyceps unilateralis (zombie-ant fungus)\|O. unilateralis]]* |
| | | | | | |
| [[#Basidiomycota (Division)]] | Agaricomycetes (Class) | Agricales | [[#Agaricaceae (Family)]] | *[[#Coprinus (Genus)\|Coprinus]]* | *[[#Coprinus comatus (shaggy ink cap mushoom)\|C. comatus]]* |
| | | [[#Polyporales (Order)]] | [[#Fomitopsidaceae (Family)]] | *[[#Fomitopsis (Genus)\|Fomitopsis]]* | *[[#Fomitopsis betulina (birch polypore mushroom)\|F. betulina]]* |
| | | | [[#Polyporaceae (Family)]] | *[[#Formes (Genus)\|Fomes]]* | *[[#Fomes fomentarius (tinder fungus)\|F. fomentarius]]* |
| | | | | | |
### Ascomyota (Division)
#### Pezizomycetes (Class)
##### Tuberaceae (Family)
##### *Tuber* (Genus)
###### truffles
Truffles are the fruiting bodies of subterranean fungi from the genus *Tuber*.[^10]
[[The aromas produced by truffles have made them among the most expensive foods in the world.]][^11]
[[Truffles with better 'chemistry' (scent) will more successfully attract animals.]][^12]
[[In many languages, the word for truffles translates to 'testicle', as in the old Castillian turmas de tierra (Earth's testicles).]][^13]
[[Truffles have long been associated with sex.]][^14]
#### Sordariomycetes (Class)
##### Hypocreales (Order)
- **See:** [[hypocrealeans]]
### Basidiomycota (Division)
#### Agaricaceae (Family)
##### *Coprinus* (Genus)
###### *Coprinus comatus* (shaggy ink cap mushoom)
[[Young, freshly-gathered shaggy ink cap mushrooms are edible. They fry well. If not eaten soon after picking, they will dye everything cooked with them black.]][^15]
[[Shaggy ink cap mushrooms (Coprinus comatus) can push their way through asphalt and lift heavy paving stones, although they themselves are delicate.]][^16]
[[Actual ink can be made from shaggy ink cap mushrooms (Coprinus comatus). They will turn to ink on their own; however, for improved ink, simply heat the ink caps with some water and cloves.]][^17]
#### Fomitopsidaceae (Family)
##### *Fomitopsis* (Genus)
###### *Fomitopsis betulina* (birch polypore mushroom)
Also known as **birch polypore mushrooms**.[^18] Have medicinal properties, known since the Copper Age (late [[Pleistocene#Neolithic|Neolithic]]).[^19]
[[Among Ötzi's belongs were wads of tinder fungus (likely used to start fire) and fragments of birch polypore mushrooms (likely medicinal).]][^20]
#### Polyporaceae (Family)
##### *Formes* (Genus)
###### *Fomes fomentarius* (tinder fungus)
This fungus has likely been used as tinder since the [[Pleistocene#Neolithic|Neolithic]].[^21]
[[Among Ötzi's belongs were wads of tinder fungus (likely used to start fire) and fragments of birch polypore mushrooms (likely medicinal).]][^22]
## fungi–human relationships
[[There are few places fungi haven't colonised. Their habitats range from the sea floor to deserts, from Antarctica to our own human guts and orifices.]][^2]
[[Robigus was the Roman god of mildew.]][^23]
### beneficial (to humans) fungi
#### agarwood
[[Agarwood is a fungal infection of Aquilaria trees.]][^24]
#### citric acid
- see: [[matter and energy#citric acid|citric acid]]
- external: [Yigitoglu, M. ‘Production of Citric Acid by Fungi’, 1992](https://jag.journalagent.com/ias/pdfs/IAS_5_2_100_106.pdf)
#### food
- see:
- [[#truffles|truffles]]
- [[#*Coprinus comatus* (shaggy ink cap mushoom)|shaggy ink cap mushrooms]]
#### industrial enzymes
[[60% of industrial enzymes are generated by fungi.]][^25]
#### ink
- see: [[#*Coprinus comatus* (shaggy ink cap mushoom)|shaggy ink cap mushrooms]]
#### medicine
[[Neanderthals made conscious use of medicinal plants and fungi—e.g. a dental abscess had been eating a type penicillin-producing mould.]][^26] (See: [[fungi#*Fomitopsis betulina* (birch polypore mushroom)|birch polypore mushroom]])
[[Australian First Nations people ‘treated wounds with moulds harvested from the shaded side of euxalyptus trees’.]][^27]
[[Ancient Egyptian papyruses (from 1500 BCE) refer to the curative properties of mould.]][^28]
[[The Talmud recounts a mould cure known as 'chamka'; chamka is produced by soaking mouldy grain in date wine.]][^29] (See: [[religion#Judaism|Judaism]])
[[In 1640, the King's herbalist in London 'described the use of moulds to treat wounds.']][^30]
##### penicillin
- [[The discovery of penicillin arguably helped shift the balance of power during World War II.]][^31]
##### other medicines
- **cyclosporine** (immunosuppressant; makes organ transplants possible)[^32]
- **cholesterol-lowering statins**[^33]
- “a host of powerful antiviral and anti-cancer compounds’[^34]
- [[life#ethyl alcohol, alcohol consumption|alcohol]]
- psilocybin
#### tinder
- see: [[#*Fomes fomentarius* (tinder fungus)|tinder fungus]]
#### waste removal
##### nuclear waste
[[One species of fungi, isolated from mining waste, is the most radiant-resistant organisms discovered. It might be capable of cleaning up nuclear waste sites.]][^35]
#### yeast
Yeasts are eukaryotic, single-celled microorganisms classified as members of the fungus kingdom. The first yeast originated hundreds of millions of years ago. At least 1,500 species are currently recognized.[^36]
The source of both [[life#ethyl alcohol, alcohol consumption|alcohols]] and [[technology#baked goods|bread]].
### fungal disease
[[There is a fungal disease, which infects amphibians, has been spread internationally via humans; it has already driven at least ninety species to extinction and threatens hundreds more.]][^37]
[[Cavendish bananas are being decimated by fungal disease and face extinction.]][^38]
### parasitic fungi
- see: [[parasites and parasitoids#parasitic fungi|parasitic fungi]]
## ↬ external resources
- [Index Fungorum](http://www.indexfungorum.org/)
+ [Mycobank](https://www.mycobank.org/)
- [Yigitoglu, M. ‘Production of Citric Acid by Fungi’, 1992](https://jag.journalagent.com/ias/pdfs/IAS_5_2_100_106.pdf)
## endnotes
[^1]: Merlin Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]* (Vintage, 2021), p. 3.
[^2]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 5.
[^3]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 2.
[^4]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 4.
[^5]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 5.
[^6]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, pp. 5–6.
[^7]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 6.
[^8]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 6.
[^9]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 7.
[^10]: ‘Truffle’, Wikipedia, last edited 12 January 2026, 13:34 (UTC), https://en.wikipedia.org/wiki/Truffle.
[^11]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 7.
[^12]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 28.
[^13]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 31.
[^14]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 31.
[^15]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 7; ‘[[Wild Food UK, ‘Shaggy Inkcap’, n.d.|Shaggy Inkcap]]’, Wild Food UK, accessed 14 January 2026, https://www.wildfooduk.com/mushroom-guide/shaggy-inkcap/.
[^16]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 7; ‘[[Wild Food UK, ‘Shaggy Inkcap’, n.d.|Shaggy Inkcap]]’, Wild Food UK, accessed 14 January 2026.
[^17]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 7; ‘[[Wild Food UK, ‘Shaggy Inkcap’, n.d.|Shaggy Inkcap]]’, Wild Food UK, accessed 14 January 2026.
[^18]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 9.
[^19]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 9.
[^20]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 9.
[^21]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 9.
[^22]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 9.
[^23]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 8.
[^24]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 32.
[^25]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 10.
[^26]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 9.
[^27]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 9.
[^28]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 9.
[^29]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 9.
[^30]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 9.
[^31]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 7.
[^32]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 10.
[^33]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 10.
[^34]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 10.
[^35]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 5.
[^36]: ‘Yeasts’, Wikipedia, last edited 12 January 2026, 14:33 (UTC), https://en.wikipedia.org/wiki/Yeast.
[^37]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 9.
[^38]: Sheldrake, *[[Sheldrake, Entangled Life, 2021|Entangled Life: How Fungi Make Our Worlds, Change Our Minds, and Shape Our Futures]]*, p. 9.